Table 1 Partition of degrees of freedom for two loci with number of alleles n₁ and n₂ (a general case)

additive (α)

2(n₁ - 1)

dominance (δ)

(n₁ - 1)²

additive (β)

2(n₂ - 1)

dominance (γ)

(n₂ - 1)²

additive × additive (αβ)

4(n₁ - 1)(n₂ - 1)

additive × dominance (αγ)

2(n₁ - 1)(n₂ - 1)²

dominance × additive (δβ)

2(n₁ - 1)²(n₂ - 1)

dominance × dominance (δγ)

(n₁ - 1)²(n₂ - 1)²

total

$n_1^2{n}_2^2$ - 1